The Dispensation for the opening of the Aylesbury and District Examining Council was issued on the 13th December 1921 (10 days before the Dispensation for the opening of the PGL was signed) The meetings where originally held on the 2nd Monday of each month, at The Red Lion Hotel, Kingsbury Square, Aylesbury, although the first meeting (Inauguration) of the Examining Council was held on Sunday 1st January 1922. The 1st President was Primo Ashdown, the 1st Vice President was Primo C. Moorecroft, the 1st Secretary was Primo T. Yates. These days Examining Council is held Bimonthly on the odd months after PGL meetings.

The issuing of the Dispensation for the opening of The Aylesbury and District Provincial Grand Lodge was approved at the December 1921 meeting of Grand Lodge in Bradford and was signed on Friday 23rd December 1921. A duplicate was issued on the 20th March 1964. The first meeting of The Aylesbury and District Provincial Grand Lodge took place in January 1922, with the 1st PGP being W. Hawkins, ROH, the 1st DPGP being H. Davis, ROH and the 1st PG Secretary being J. Francombe CP

The Dispensation for the opening of The Aylesbury Knights Chapter No 209 was issued on Wednesday 30th May 1934, a duplicate was issued on the 16th November 1987. The first meeting of the Aylesbury Knights Chapter No 209 was on Saturday 16th June 1934. At one stage the Aylesbury and District Province had two Knights Chapter. The Aylesbury Knights Chapter and The Claydons Knights Chapter. The Claydons Knights Chapter was formed by members of The Province who lived to the North and West of the Aylesbury and District Province who could not get into Aylesbury due to the lack of their own and or public transport. Their meetings where held in Buckingham

The Dispensation for the opening of the Triumph Lodge No 7207 was signed on Saturday 20th January 1934. The founder members of the Triumph Lodge were Brothers, T. Turner, CP, S. K. Hutchings, CP, B. Cross, CP and C. Henley, CP.

The Dispensation for the opening of the Conformity Lodge No 7546 was signed on Thursday 20th March 1941. Unfortunately the Inauguration date has worn off of the Warrant, this would have been on either the 13th or 20th April 1941. The name Conformity was chosen as the lodge was opened to conform to the hours worked by the people in the utility services who where exempt from war duty so that they could go to a lodge either before starting or after finishing work on a Sunday. The founder members of the Conformity Lodge were Brothers: C. W. J. Poulton, CP, H. J. Burchell, CP, A. Williamson, CP and J. Goss, KOM. The longest serving member of the Conformity Lodge to date is V. I. Clarke, ROH, who joined the order on Sunday 21st September 1947 at the age of 21

The Dispensation for the opening of the RAF Halton Lodge No 7622 was signed on Thursday 22nd July 1943. The Inauguration of the Lodge took place on Monday 9th August 1943. The Founding members of the RAF Halton Lodge were Brothers: J. D. Paley, CP, J. H. McKeone, KOM, W. L. Shears, ROH, and W. Moore, ROH.

The Dispensation for the opening of the Waddesdon Lodge No 8012 was signed on Wednesday 22nd September 1948. The Lodge was Inaugurated on Friday 8th October 1948. The Founder members of The Waddesdon Lodge were Brothers: W. Hollaway, CP, J. Truman, CP, J. E. Carter, CP and C. W. J. Poulton, ROH. The Lodge was named after Waddesdon Village a few miles West of Aylesbury on the A41.

The Dispensation for the opening of the Oving Lodge No 8022 was signed on Thursday 7th October 1948. The Lodge was Inaugurated on Saturday 13th November 1948. The Founder members of The Oving Lodge were Brothers: W. G. R. McAndrew, ROH, G. C. Peters, KOM, A. G. Dobbs, CP and B. J. H. Fowler, CP. The first home of The Oving Lodge was The Thistle PH in Whitchurch a small village a few miles West of Aylesbury on the A413. The Oving Lodge was named after OVING Village an even smaller village than Whitchurch.

The Province Boundary changed in March 1959 when Wing Village was transferred from The North Bucks Province.

The Victor Lacey Lodge No 8867 was named after The Victor Bomber and a Village called Lacey Green not far from Princes Risborough where the Lodge had its first home.

The Dispensation for the opening of the Hearts of Oak Lodge No 9729 was signed on Wednesday 18th March 1970. The Lodge was inaugurated on Wednesday 22nd April 1970. The Founder members of the Hearts of Oak Lodge were Brothers: G. J. Wright, ROH, C. R. Heydon, CP, R. B. Guinard, KOM and R. Day, CP.

The Vale Lodge No 7032 Closed at the end of 1970.